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Infraorder Brachyura // Family Polybiidae // Genus Liocarcinus
  Liocarcinus vernalis (Risso, 1827) Larvae development
Synonyms: Portunus dubius nova species? Rathke, 1837 Only for the Black Sea
Portunus holsatus Fabr. Markuzen, 1868 Only for the Black Sea
Macropipus holsatus (Fabricius, 1798) Băcescu, 1967 Only for the Black Sea
Polybius vernalis (Risso, 1827) Micu&Micu, 2006 Only for the Black Sea
Macropipus barbarus (Lucas, 1846) WoRMS, 2017  
Macropipus vernalis (Risso, 1827) WoRMS, 2017  
Polybius (Polybius) vernalis (Risso, 1827) WoRMS, 2017  
Portunus barbarus Lucas, 1846 WoRMS, 2017  
Portunus valentieni Cocco, 1833 WoRMS, 2017  
Portunus vernalis Risso, 1827 WoRMS, 2017  
It is found in the northeastern Atlantic from the coast of Norway to Portugal, as well as in the seas of the Mediterranean basin and in the Black Sea (Kobyakova, Dolgopolskaya, 1969).
Numerous species, widespread in all regions of the Black Sea to a depth of 40 m. Prefers sandy soil with shells, but can also be found on silted sand among thickets of zostera. Usually, these crabs actively move on the surface of the ground and, in case of danger, quickly burrow into the sand. Females with eggs on the surface of the sand during daylight hours are not found, but at night they can be found in large numbers within this biotope. In the different places of habitat can be have different colors of individuals.
North East Atlantic Black Sea    
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